The wifi and the LAN port are bridged. This means packets from one can flow to and from the other. This does not mean they will mind you. Above bridging is IP address and subnetting. To traverse the bridge you have to be on the same IP network.

When the destination is the same subnet, your computer asks everyone "hey who has IP address 172.21.13.153, and the device sees this and says I am at mac address … Because you have a bridge, and change your IP address, this is why traffic flows over the bridge to the device on the switch.

When the destination is a different subnet your computer says that is not local, I’ll send that to my default gateway. So when you are set to DHCP, your computer forwards traffic for 172.21.13.153 to you wireless internet connection. Not gonna find it there.

To let the AR300 sort things out, I am guessing we will need to define 2 additional networks , not sure from your wireshark above of the topology.

Can you get to Luci (on a MT300N-V2 its Other Settings->Advanced)? If so we might be able to sort it out. I think you will need to plug one of the switches into the wan port though, and we can repurpose its use, I hope without upsetting the GL.inet interface.

Your use case seems somewhat removed for what the AR300M was designed for!

This means what you are trying to do is fairly complicated.

I have not got an AR300M, so I do not know how they are configured and use openwrt, but what you want to do is quite possible in openwrt.

Simon

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